U.S. Fish & Wildlife Service
Status and Harvests of
Sandhill Cranes 2023
Mid-continent, Rocky Mountain,
Lower Colorado River Valley and
Eastern Populations
Acknowledgments
This report provides population status, recruitment indices, harvest trends, and other management
information for the Mid-Continent (MCP), Rocky Mountain (RMP), Lower Colorado River Valley (LCRVP),
and Eastern (EP) populations of sandhill cranes. Information was compiled with the assistance of many
biologists from across North America. We acknowledge the contributions of: P.P. Thorpe, T. Liddick, J.
Drahota, and D. Fronczak for conducting annual aerial population surveys; D. Collins and R. Vanausdall
for conducting the RMP productivity survey; K. Fleming for conducting the U.S. Federal harvest surveys for
the MCP; S. Olson and L. Harding for compiling population and harvest information collected on sandhill
cranes in the Pacific Flyway; R. Pierce and D. Fronczak for compiling population information for the EP;
and D.S. Benning, R.C. Drewien, J.A. Dubovsky, and D.E. Sharp for their career-long commitment to
sandhill crane management. We especially want to recognize the support of the state and provincial
biologists in the Central, Pacific, and Mississippi Flyways for the coordination of sandhill crane hunting
programs and especially the distribution of crane hunting permits and assistance in conducting annual
cooperative surveys and sharing harvest data. Thanks to T. Cooper and D. Collins for reviewing earlier
drafts of this report.
Citation: Seamans, M.E. 2023. Status and harvests of sandhill cranes: Mid-Continent, Rocky Mountain,
Lower Colorado River Valley, and Eastern Populations. Administrative Report, U.S. Fish and Wildlife
Service, Lakewood, Colorado. 17pp + tables and figures.
This report contains annual estimates of migratory bird abundance, harvest, and hunter participation and
activity. Due to the large volume of data, the number of years, and geographic areas involved, data tables
may be large and complex. Readers that may need help reading and interpreting the data, or that may
need data presented in an alternative format to facilitate reading and interpretation, should contact the
author at mark_seam[email protected].
Division of Migratory Bird Management sandhill crane reports are available online at
https://www.fws.gov/library/collections/sandhill-crane-population-status-reports
18
STATUS AND HARVESTS
OF
SANDHILL CRANES
MID-CONTINENT, ROCKY MOUNTAIN, LOWER COLORADO
RIVER VALLEY and EASTERN POPULATIONS
2023
Mark E. Seamans, Division of Migratory Bird Management, U.S. Fish and Wildlife Service,
Lakewood, Colorado.
Abstract: The U.S. Fish and Wildlife Service, working with partners, annually assesses the
population status and harvest of four populations of sandhill cranes: the Mid-continent, Rocky
Mountain, Lower Colorado River Valley, and Eastern populations. Annual abundance estimates
of the Mid-Continent Population (MCP) of sandhill cranes were relatively stable from 1982 to the
mid-2000s. Some of the MCP annual indices have increased in recent years and are more
variable interannually compared to historic values. The spring 2023 estimate of abundance for
sandhill cranes in the Central Platte River Valley (CPRV), Nebraska, corrected for visibility bias,
was 1,259,199 birds. This estimate is 83% higher than the 2022 estimate and is a record high for
the MCP. The photo-corrected average using the most recent 3 years of data (2011–23) is
909,046, which is well above the established population-objective range of 350,000475,000
cranes. All Central Flyway states, except Nebraska, allowed crane hunting in portions of their
states during 2022–23. An estimated 23,529 Central Flyway hunters participated in these
seasons. This estimate was 12% higher than the number that participated in the previous season
for the same states. Hunters harvested 56,178 MCP cranes in the U.S. portion of the Central
Flyway during the 2022–23 season. The fall 2022 pre-migration survey for the Rocky Mountain
Population (RMP) counted 18,632 cranes, 22% lower than the count from 2021. The 3-year
average was 22,744 sandhill cranes, which exceeds the established population objective of
17,00021,000 for the RMP. Hunting seasons during 2022–23 in portions of Arizona, Idaho,
Montana, New Mexico, Utah, and Wyoming resulted in a harvest of 1,586 RMP cranes, an 11%
increase from the previous year’s harvest. The Lower Colorado River Valley Population (LCRVP)
survey results indicated a 25% increase from 2022 (3,787 birds) to 2023 (4,719 birds). The 3-
year average is 4,796 LCRVP cranes, which is above the population objective of 2,500 birds. The
Eastern Population (EP) sandhill crane fall survey index for 2022 (107,164) was a 19% increase
from the previous year, and well above the objective of 30,000 cranes for this population. A total
of 1,085 cranes were harvested in Alabama, Kentucky, and Tennessee during the 2022–23
season, which was 30% higher than last year.
19
Introduction
The MCP of sandhill cranes, numerically the most abundant of all North American crane
populations, is comprised of lesser (Antigone canadensis canadensis) and greater (A. c. tabida)
subspecies of sandhill cranes. The breeding range extends from northwestern Minnesota,
northern Ontario, and western Quebec, then northwest through Arctic Canada, Alaska, and into
eastern Siberia. The MCP wintering range includes western Oklahoma, New Mexico,
southeastern Arizona, Texas, and northern portions of Mexico (Fig. 1). Extensive spring aerial
surveys on major concentration areas that are corrected for observer visibility bias provide annual
indices of abundance that are used to estimate population trends. These surveys are conducted
in late March, at a time when cranes that wintered in Mexico, Arizona, New Mexico, and Texas
usually have migrated northward to spring staging areas along the Platte River in Nebraska, but
before spring "break-up" conditions allow cranes to move into Canada (Benning and Johnson
1987). The MCP Cooperative Flyway Management Plan (Central, Mississippi and Pacific Flyway
Councils 2018) established regulatory thresholds for changing harvest regulations that are based
on an objective of maintaining sandhill crane abundances at 19822005 levels (i.e., spring index
of 349,000–472,000 [̅ = 411,000 ± 15%]), rounded up to the nearest 5,000 birds. Sandhill crane
hunters are required to obtain either a sandhill Crane hunting permit and/or register under the
Harvest Information Program (HIP) to hunt MCP cranes in the U.S. portion of the Central Flyway,
Minnesota in the Mississippi Flyway, and Alaska in the Pacific Flyway. The permits or HIP
registration records provide the sampling frame to conduct annual harvest surveys. In Canada,
the harvest survey is based on the sales of Federal Migratory Bird Hunting Permits, which are
required for all crane hunters.
The RMP is comprised exclusively of greater sandhill cranes that breed in isolated river valleys,
marshes, and meadows of the U.S. portions of the Central and Pacific Flyways (Drewien and
Bizeau 1974). The highest nesting concentrations are in western Montana and Wyoming, eastern
Idaho, Colorado, and Utah. The RMP migrates through the San Luis Valley (SLV) in Colorado
and winters primarily in the Middle Rio Grande Valley, New Mexico, with smaller numbers
wintering in the southwestern part of New Mexico, in southeastern Arizona, and at several
locations (~14) in the Northern Highlands of Mexico (Fig. 2). During 198496, the RMP was
monitored at spring stopover areas in the SLV. However, cranes from the MCP also began to
use this area, which confounded estimates of RMP abundance. In 1995, a fall pre-migration
(September) survey replaced the spring count as the primary tool for monitoring population
change. The RMP Cooperative Flyway Management Plan established a population objective
(17,00021,000 birds), and identifies surveys used to monitor recruitment and harvest levels that
are designed to maintain a stable abundance (Pacific Flyway Council and Central Flyway Council
2016). The plan contains a formula for calculating allowable annual harvests consistent with the
goal of staying within the range of the population objective. All sandhill crane hunters in the range
of the RMP must obtain a state permit to hunt cranes, which provides the sampling frame for
independent harvest estimates and allows for assignment of harvest quotas by state. In many
areas, harvest estimates are supplemented by periodic mandatory check-station reporting.
The LCRVP is numerically the least abundant of the six migratory populations of sandhill cranes
recognized in the U.S. (Drewien et al. 1976, Drewien and Lewis 1987). The LCRVP is comprised
exclusively of greater sandhill cranes that breed primarily in northeastern Nevada and southwest
Idaho, with smaller numbers in parts Utah (Fig. 3), and winters largely in the Colorado River Valley
of Arizona and Imperial Valley of California (Grisham et al. 2018). LCRVP cranes are thought to
have the lowest recruitment rate of any sandhill crane population in North America (Drewien et
20
al. 1995). In the fall, these cranes leave breeding areas during late September-early October,
congregate at several staging areas, and migrate through eastern Nevada to wintering areas.
Wintering areas historically extended south along the Colorado River to near its delta with the
Gulf of California. However, the current wintering distribution is concentrated at Cibola National
Wildlife Refuge, on areas just north of the Cibola National Wildlife Refuge belonging to the
Colorado River Indian Tribes in southwestern Arizona, areas within and near the Sonny Bono
Salton Sea NWR in southern California, and the Gila River in Arizona. Collectively, these areas
are believed to winter more than 90% of the total cranes in the LCRVP. Spring migration is
generally initiated as early as the first week of February. Since 1998, an aerial cruise survey has
been conducted that covers the four main winter concentration areas.
The EP, which consists of greater sandhill cranes, has rebounded from near extirpation in the late
1800’s (Walkinshaw 1949, 1973; Leopold 1949). Management actions, such as regulating take
and the protection and restoration of habitat, allowed this population to increase to a level that
exceeded 30,000 cranes by 1996 (Meine and Archibald 1996). Most EP cranes breed across the
Great Lakes region (Wisconsin, Michigan, Ontario, and Minnesota); however, the range of this
population is currently expanding in all directions (Fig. 4) (Lacy et al. 2015). By early fall, EP
cranes leave their breeding grounds and congregate in large flocks on traditional staging areas
throughout the breeding range. During migration, EP cranes use traditional stopover areas which
include Jasper-Pulaski Fish and Wildlife Area in northwest Indiana and Hiawassee State Wildlife
Refuge in southeast Tennessee. Historically, EP cranes primarily wintered in southern Georgia
and throughout Florida (Walkinshaw 1973, Lewis 1977, Tacha et al. 1992, Meine and Archibald
1996). Recent annual Midwinter Survey data, conducted by state and federal agencies, show
substantial numbers of cranes wintering farther north into Kentucky, Tennessee, and even Indiana
in some years (2013–2020 U.S. Fish and Wildlife Service [unpublished data], Fronczak et al.
2017, Urbanek 2018).
Mid-Continent Population of sandhill Cranes
Sport hunting seasons for MCP cranes were not allowed in the U.S. 1918–60. In the Central
Flyway, areas open to hunting were gradually expanded during 196174, but since that time have
remained relatively stable. Operational hunting seasons are now held annually in portions of
Colorado, Kansas, Montana, New Mexico, North Dakota, Oklahoma, South Dakota, Texas, and
Wyoming. Nebraska is the only Central Flyway state that does not have a sandhill crane sport
hunting season. Areas open to crane hunting in the Central Flyway during 2021–22 are shown
in Fig. 5. Beginning in 2010, Minnesota, a Mississippi Flyway state, opened a limited hunt in the
northwest portion of the state.
During 196174, hunters gradually improved their knowledge of sandhill cranes and improved
their hunting success. During 197585, a tradition of sandhill crane hunting became established.
Together with improvements in equipment (decoys, calls, clothing, blinds, etc.) and a shift from
pass-shooting and hunting on roosts to decoy-hunting in fields, crane hunter success increased
(Sharp and Vogel 1992). Dubovsky and Araya (2008) found that in the late 1990s and early 2000s
hunters were more successful in harvesting 2 or 3 cranes per day than they were during the early
1980s. Average seasonal bags declined in the Central Flyway during the late 1990s and early
2000s, but during the last several seasons have increased to levels observed in the late 1980s to
late 1990s (Fig. 12).
21
For most states, sandhill crane seasons began in relatively small areas, and expanded
incrementally in subsequent years as experience with the seasons was gained. For example,
sandhill crane seasons in North Dakota resumed in 1968 after being closed following the signing
of the Migratory Bird Treaty Act in 1918. During 196879, the number of counties open for crane
hunting increased from 2 to 8, and increased to 30 during 198092 and were grouped into two
zones that were west of Highway 281. Beginning in 1993, the zones were eliminated, and Federal
frameworks were fully utilized for the designated hunting area (Sharp and Cornely 1997). In 2001,
designated hunt areas in North Dakota and Texas were expanded, with the new areas having
reduced frameworks of 37 days compared to 58 in other areas and a reduced daily bag. In 2014,
North Dakota increased season length in the eastern zone to 58 days but kept the 2-bird daily
bag limit; harvest data suggested there would be negligible effects on that segment of the
population. Kansas was the most recent Central Flyway state to initiate a crane hunting season
in 1993. Initially, crane hunting was open only in portions of 17 counties, but by 2003 the area
was expanded to 62 counties, essentially the entire western portion of the state (Sharp et al.
2010). Also, during early years of these seasons, bag limits and shooting hours often were more
restrictive than Federal frameworks allowed. Beginning in the 201920 season, South Dakota
moved their hunt boundary eastward from where it had been historically.
MCP harvest areas have remained relatively consistent from year to year; however, the levels of
harvest vary with respect to many factors including changes in hunting pressure, land use, and
environmental factors. Most shifts in annual harvests occur locally, but large-scale changes in
harvest distributions also have occurred. Since the late 1990s, the annual harvest has generally
increased declined in North Dakota and, until recently, increased in Saskatchewan (Tables 5A
and 6). Causal factors for these changes have not been determined but are likely different
because birds staging in Saskatchewan are largely from the West-central Canada-Alaska
breeding affiliation whereas those in North Dakota are from the East-central Canada-Minnesota
breeding affiliation (Krapu et al. 2011). Increased hunting pressure in Saskatchewan, mainly by
non-resident U.S. hunters (Araya et al. 2010), has likely contributed to increases in harvests
whereas declines in harvests in North Dakota appear to be more complex and involve several
interrelated factors, likely including changes in hunting pressure, land-use changes, and
environmental conditions. More recently, in Texas the interest in crane hunting has increased,
with the number of hunters and annual harvest approximately doubling from 2016 to 2021 (Table
3A).
The MCP included at least 510,000 sandhill cranes in March 1982, the last extensive survey
involving high-altitude vertical photography of major spring migration staging concentrations.
Beginning in 1982, an intensive photo-corrected ocular-transect survey of Nebraska's Central
Platte River Valley (CPRV) and ocular assessments from other spring staging areas have been
used to monitor the annual status and trends for this population (Table 1). Use of the CPRV count
in the development of annual harvest recommendations relies on the premise that a high
proportion (>90%) of the MCP are in the CPRV at the time of the annual survey. Recent research
with radio-tracked birds suggests that the proportion of MCP cranes in the CPRV during the
survey varies by year (Pearse et al. 2015). Annual variability in weather patterns can reduce the
percentage below 90% in some years. However, conducting the survey a few days earlier or a
few days later likely would not result in a 'better' count (i.e., a higher proportion of birds being in
the CPRV), because birds migrate into and out of the area continuously (Pearse et al. 2015).
The count from the March 2023 survey for MCP was 1,259,199 cranes (Table 1, Fig. 6) in the
CPRV, which represents a record high estimate since the survey was initiated in 1974 (Liddick
2023). The natural log-transformed annual photo-corrected estimates for the CPRV portion of the
22
survey suggest an increasing population trend (P = 0.01) since 2006 with higher counts in several
of the recent surveys (Fig. 7); however, estimates also have more interannual variability in recent
years relative to historic values, resulting in a poor fit of the trend to the data (R
2
= 0.36). The 3-
year-average index for photo-corrected estimates in the CPRV during the most recent three years
that surveys have been conducted (2021-23) was 909,046 cranes, which is well above the
management objective level (350,000-475,000) for this population (Fig. 8). Due to COVID
restrictions the survey was not completed in 2020, and ground counts used to correct aerial counts
were not done in 2021.
Since 1975, special sandhill Crane Hunting Permits, or more recently HIP certification, have been
required for crane hunters participating in seasons in the Central Flyway. Additionally, a limited
MCP sandhill crane hunt was offered in Minnesota starting in 2010, for which a state-issued permit
is required for hunters to participate. A sample of these permittees is mailed questionnaires soon
after the completion of each hunting season. The resulting responses enable estimation of
hunting activities and success (Martin 2007). Estimated numbers of hunters registering as
sandhill crane hunters in Texas has been increasing since 1997 when crane hunting was included
in the combination licenses issued by the state, with a record high of 122,553 permits issued in
2008. In 2009, Texas revised their licensing system and crane hunters now must go to selected
locations to obtain their permit, which resulted in a 91% decrease in the number of permits issued
to individuals in 2009 compared to 2008. Thus, the number of crane hunters in Texas likely did
not decrease as suggested by the number of permits issued; rather, the number of hunters
classified as crane hunters by the Texas registration process declined. For the 2019-20 season,
Oklahoma did not provide information needed to estimate hunter activity and harvest in time to
conduct surveys of their hunters; thus, no estimates of the number of hunters or their harvest of
cranes is available. For the 202021, 202122, and 202223 seasons, Oklahoma reported the
number of permits as they did prior to 2019, with the addition of hunters who received a free online
crane hunting permit (Table 2A). These free permits were not previously in the HIP sample of
Oklahoma crane hunters, and their inclusion resulted in a large increase in the number of hunters
for these seasons. The number of crane hunters in Texas increased 89% from 2019 to 2020
(Table 3A), and appears to be the result of crane hunting becoming more popular. During the
2022–23 season in the Central Flyway, 87,577 hunters were either HIP-certified or obtained crane
hunting permits, which were not limited in number (Tables 2A, 2B), with 23,529 of these
individuals hunting at least one time (Tables 3A, 3B, Fig. 9). The 202223 number of active
hunters in the Central Flyway increased 12% from the previous year (Fig. 9). During 2022–23,
the number of hunters in Texas comprised 71% of the sandhill crane hunters in the Central
Flyway. Minnesota issued 1,424 permits for the 202223 season and had 898 active hunters (4%
decrease and 7% increase, respectively, from the 2021–22 season).
Federal frameworks for most areas in the Central Flyway allow daily bag/possession limits of 3/9,
which most states selected. Portions of North Dakota, Texas and Minnesota have had lower bag
and possession limits of 2/6; the bag/possession limit in Minnesota was lowered to 1/3 beginning
with the 2018-19 season. Specific dates selected by states in the Central Flyway and Minnesota
for 2022-23 were similar to those of previous hunting seasons (Table 4).
An index to crippling-loss rate (number of cranes lost/[number of cranes lost + retrieved]) in the
U.S. portion of the Central Flyway has declined (R
2
= 0.91, P < 0.01) from over 16% in 1975 to a
preliminary estimate of 1.1% during the most recent hunting season (Fig. 10). During the 2022
23 season the number of days afield per hunter (2.18) was 9% lower than that of the previous
year (Fig. 11) and was 28% lower than the long-term average of 3.02. The preliminary estimate
of seasonal bag per hunter was 2.39 birds (Fig. 12), which is 8% higher than the long-term
23
average of 2.22. The preliminary estimate of retrieved and unretrieved mortality associated with
the sport harvest in the Central Flyway was 8% lower than the previous year's estimate (Fig. 13).
The increasing trend (R
2
= 0.71, P < 0.01) in the Central Flyway’s harvest of MCP cranes during
19752022 likely is related to improved knowledge of crane behavior, hunting techniques, and
hunter success (Sharp and Vogel 1992, Dubovsky and Araya 2008), and increased numbers of
cranes available for harvest in recent years due to growth in the MCP.
Cranes from the MCP also occur in the RMP hunt areas in Arizona, New Mexico, Alaska (Tables
5A, 5B), Canada, and Mexico. Estimates for the 2022–23 sport harvest in Canada (Alberta,
Manitoba, and Saskatchewan) were not available at the time this report was completed; historic
estimates are provided in Table 6. For Alaska, sandhill cranes harvested in Game Management
Units (GMUs) 11-13 and 18-26 are believed to be MCP cranes, while cranes harvested in GMUs
1-10 and 14-17 are believed to be Pacific Coast Population cranes. There also is some
intermingling of MCP cranes with RMP cranes in portions of New Mexico and Arizona; however,
periodic bag checks allow estimates of harvests for each population. The estimated MCP harvest
for the RMP hunt areas in Arizona, New Mexico, and Alaska combined was 1,799 cranes for
2022–23. In the 13th year of Minnesota’s sandhill crane hunt the harvest (863 cranes) increased
by 53% from the previous year. No annual harvest surveys are conducted in Mexico, but annual
MCP harvests probably are <10% of the retrieved harvest in the U.S. and Canada (R. Drewien
and D. Nieman, personal communication). This assumed low level of harvest was supported by
an independent assessment of harvest in Mexico (Kramer et al. 1995). Because harvest
estimates for Canada were not available, the 2022–23 estimates of retrieved and unretrieved kill
of MCP cranes by sport hunters throughout their range was not calculated. Historic information is
provided in Tables 7A and 7B, and Fig. 14.
To assess the relative rates of change between population size (abundance) and harvest, we
periodically assess trends in these parameters. We used linear regression on the natural log-
transformed values for these variables for the years 1982-2021. Because >10% of the MCP
occurs outside the CPRV in the spring of some years, we combined the photo-corrected counts
in the CPRV with the ocular cruise estimates from areas outside the CPRV for analyses of
population abundance. For harvest, we used only the estimates of retrieved harvest for the
Central Flyway, Minnesota, and MCP cranes harvested in hunt areas in Arizona and New Mexico,
Alaska, and Canada, because crippling-loss rates for the latter three areas are unknown and there
are no empirical estimates of harvest from Mexico. Regression of the log-transformed values
indicate a significant slope for the abundance values (P < 0.01; R
2
= 0.36; slope = +1.6% per year
change), suggesting an increasing trend in the abundance of cranes over the time frame. The
regression of the harvest values also indicates an increase in the rate of harvest over that same
time (P < 0.01; R
2
= 0.71; slope = +2.3% per year) (Fig. 15). These results suggest that the
annual change in harvest is increasing faster than the rate of growth in crane abundance.
Subsistence harvest levels of MCP sandhill cranes historically were poorly documented.
However, the 1997 U.S./Canada Migratory Bird Treaty Amendment identified improvements that
should be made to sandhill crane harvest-monitoring programs in both the U.S. and Canada.
Harvest surveys conducted during 20062017 on the Yukon-Kuskokwim (Y-K) Delta, Alaska,
reported an average MCP harvest of 2,896 adults and fledged young and an average of 1,183
eggs (data from Naves and Keating 2019). The harvest estimate for birds is relatively similar to
the 1985-2005 average (Wentworth 2007) of 3,151 adults and fledged young taken by
subsistence hunters on the Y-K Delta, but that for eggs is 124% higher than the 1985-2005
average of 528 eggs. Efforts are being made to gather additional information on subsistence
harvests for the remainder of Alaska, Siberia, and Canada.
24
Rocky Mountain Population of Greater sandhill Cranes
The RMP was not hunted in the U.S. from 1918–80. Arizona initiated the first modern-day season
in 1981. Since that time hunting programs have been guided by a cooperative management plan,
including a harvest strategy that has been periodically updated and endorsed by the Central and
Pacific Flyways (Kruse et al. 2008). The harvest strategy for the RMP calculates an allowable
harvest based on crane survey counts and recruitment relative to the population objective. Thus,
allowable harvest changes annually based on the status of the birds.
Counts conducted in the SLV during the spring migration suggested that the number of RMP
cranes was relatively stable during 198496 (Table 8). However, survey biologists found that
these estimates contained increasing numbers of the MCP (lesser subspecies). An adjustment,
using ground-derived proportions, was made to correct for the lesser subspecies but was not a
viable approach for the long-term (Benning et al. 1996). In 1996, the survey was discontinued
(Table 8). In 1997, an attempt was made to survey these cranes during the fall (October) in the
SLV, but MCP cranes also were present at that time. Biologists concluded that neither a spring
nor a fall count in the SLV would result in a reliable index to the abundance of the RMP. As an
alternative, a cooperative 5-state September pre-migration staging-area survey, experimentally
tested in 1987 and 1992, has been ongoing operationally since 1995. Because there appears to
be minimal commingling of RMP cranes with cranes from other populations during that time, the
September pre-migration survey for the RMP appears to be a good alternative to either a spring
or fall survey in the SLV and was designated as the official count for the RMP in 1997 (Table 9).
Although operational in 1995 and 1996, the survey was variable in timing and survey effort. What
appears to be lower population estimates (Table 9) in 1995 and 1996 is likely more an artifact of
inconsistent survey effort (R. Drewien, personal communication).
The Cooperative Flyway Management Plan (Pacific Flyway Council and Central Flyway Council
2016) recommends using the most recent three-year running average of the September survey
to determine status of the RMP. The 2022 September pre-migration survey resulted in 18,632
cranes counted, a 22% decrease from the count in 2021 (Thorpe et al. 2022) (Table 9). The 3-
year average is 22,744 which is 4% lower than the previous 3-year average (23,630) and exceeds
the range of the established population objective (17,00021,000) (Fig. 18).
During 198695, important breeding areas in the Intermountain West experienced extremely dry
conditions and indices of recruitment (% juveniles) were low (generally between 46%) (Fig. 19).
A return to more favorable breeding conditions during 199699 resulted in higher recruitment
rates (812%), but drier conditions resulted in lower production during 2000-02. Since 2003
recruitment rates generally have been above-average due to improved wetland habitats and
favorable spring and summer breeding conditions. The recruitment rate of 10.8% (30% above
the long-term [1972–2021] average of 8.2) suggested above average nesting conditions, but a
mean brood size of 1.08 (Collins and Vanausdall 2022) indicated the opposite in 2022.
Special limited hunting seasons during 2022–23 resulted in a harvest of 1,586 RMP sandhill
cranes (Tables 10A, 10B), which was 11% greater than last year (Fig. 16). Based on the surveys
conducted last fall which resulted in 3-year (2020–22) average values that were lower for
abundance but higher for recruitment (Figs. 18, 19), management guidelines allow for a maximum
allowable take of 2,543 birds during the 2023-24 hunting season, an 8% decrease from that for
the 2022–23 season.
25
Lower Colorado River Valley Population of Greater sandhill Cranes
The LCRVP is the smallest of the migratory populations of sandhill cranes in North America. The
range of this population is believed to overlap ranges with the Rocky Mountain and Central Valley
populations. Historically, winter counts of the LCRVP were not well-coordinated or conducted
using a consistent methodology. However, efforts have been made to standardize areas
surveyed and the timing of the survey to obtain more accurate counts and increased ability to
determine trends in population abundance. Beginning in 1998, a coordinated winter aerial cruise
survey with a fixed-wing aircraft has been conducted at the four major wintering areas: Cibola
NWR, Colorado River Indian Tribal lands, Sonny Bono Salton Sea NWR, and the Gila River.
Collectively, these counts are believed to contain >90% of the total number of cranes in this
population. The counts are not corrected for cranes present but not seen by aerial crews, and
therefore have unknown bias and precision. The fixed-wing aircraft survey was again conducted
in 2023. The survey resulted in a count of 4,719 birds in 2023, a 25% increase from the previous
years count of 3,787 birds (Table 11, Fig. 20). The current 3-year average for the winter count
is 4,796 cranes.
The LCRVP was not hunted after the signing of the Migratory Bird Treaty Act in 1918. In 2007,
the Service completed an Environmental Assessment entitled “Proposed hunting regulations for
the Lower Colorado River Valley Population of Greater sandhill Cranes in the Pacific Flyway”
(U.S.D.I. 2007). In 2008, the Service determined that a small allowable harvest (about 30) could
be allowed on this population in years when the 3-year average of winter counts exceeded 2,500.
The hunting season is guided by a cooperative management plan (Pacific Flyway Council 1995)
which includes methodology for determining allowable harvests and allocation of the harvest.
Once a hunting season is initiated, this season would be experimental for 3 years. After the 3
years, the season would be reviewed and revised if necessary.
A limited youth hunting season for this population was conducted during 2010 in Arizona, the only
state that has hunted these cranes. No LCRVP cranes were harvested. The Pacific Flyway
currently has no plans to conduct hunts for LCRVP cranes.
Eastern Population of Greater sandhill Cranes
In 1979, the U.S. Fish and Wildlife Service initiated a coordinated fall index survey of historic EP
migratory staging areas in the Mississippi and Atlantic Flyways. This survey is conducted
annually in late October by volunteers and agency personnel who count the number of cranes at
staging areas throughout the EP range. Overall, the survey documented a long-term increasing
trend in EP cranes with an average growth rate in the population of 3.9% per year (1979-2009)
(Amundson and Johnson 2010). A more recent analysis indicates the growth rate has increased
to 4.4% per year (U.S. Fish and Wildlife Service, unpublished data). The most recent fall count
from 2022 was 107,164, which was 19% higher than the 2021 index of 90,029. The 3-year
average is 97,385 (Table 12, Fig. 21). This index is not a statistically designed population
estimate; however, the index does reasonably represent a conservative population estimate for
EP cranes.
In 2010, the Mississippi and Atlantic Flyway Councils endorsed a management plan for EP cranes
(Ad Hoc Eastern Population sandhill Crane Committee 2010). One of the plan’s provisions
26
included guidelines for potential harvest of this population when the 3-year average of the fall
survey is above 30,000 cranes. Kentucky and Tennessee initiated experimental hunting seasons
in 2011 and 2013, respectively; the season in Kentucky became operational in 2015 and that for
Tennessee in 2017. Alabama initiated an experimental season beginning in the 2019-20 season
(Table 13), and the season will became operational in the 2023-24 season. Hunting seasons for
this population of sandhill cranes are allowed between September 1 and January 31 and have a
maximum length of 60 days. Actual season dates have been from early-December to late-January
in Kentucky and late November to late January in Tennessee (Table 13). During recent years the
seasons in each state have extended from early December through the end of January. According
to the hunt plan, the number of tags a state can issue cannot exceed 10% of the state’s five-year
average peak crane abundance. Each tag allows a hunter to harvest one crane. Hunters in all
three states are required to complete mandatory crane identification training, tag and report
harvested birds, and complete a post-season survey. In the 2022–23 season, Kentucky issued
1,148 tags to hunters, who harvested 180 cranes (Table 14)(J. Brunjes, Kentucky Department of
Fish and Wildlife Resources, personal communication). Tennessee issued 2,500 tags to hunters
and 640 cranes were harvested in 202223 (Table 14) (J. Feddersen, Tennessee Wildlife
Resources Agency, personal communication). Alabama issued 1,200 tags to hunters in 2022-23
and 265 cranes were harvest harvested (Maddox 2023) (Table 14). The total number of Eastern
Population sandhill cranes harvested during 2022–23 hunting season was 1,085 birds, which was
30% higher than last year.
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29
Table 1. Annual spring abundance indices for the Mid-Continent Population of sandhill cranes. Estimates presented for: (1) the Central Platte River
Valley (CPRV), NE; (2) other areas in Nebraska outside the CPRV and in other states; and (3) total combined estimates.
Year
CPRV
Ocular
Cruise
Transect
CPRV
Ocular
Transect
CPRV
Photo
Corrected
Ocular
Transect
Annual
CPRV
Photo
Corrected
Ocular
Transect
3-Year AVG
Other
KS
CO
a
OK
a,b
NM
a
WY
b
Total
Ocular
Cruise
Transect
Total
Ocular
Transect
Total Photo
Corrected
Ocular
Transect
Annual
Total Photo
Corrected
Ocular
Transect
3-Year AVG
1974
162,600
1,900
0
400
0
177,100
1975
223,600
900
500
100
100
227,500
1976
147,500
300
0
100
1,000
152,500
1977
173,400
1,600
0
400
12,500
220,000
1978
149,800
188,582
700
0
0
2,300
159,900
198,682
1979
203,574
1,100
500
1,500
0
209,274
1980
223,400
254,417
4,100
0
100
500
234,500
265,517
1981
248,882
11,200
500
0
0
290,682
1982
347,996
417,263
2,000
2,800
0
100
367,796
437,063
1983
306,316
343,378
200
0
200
tr
317,816
354,878
1984
222,710
261,802
340,814
900
0
1,100
tr
243,610
282,702
358,214
1985
378,127
514,763
373,314
3,000
393,827
530,463
389,348
1986
317,025
353,040
376,535
200
320,325
356,340
389,835
1987
383,581
416,058
427,954
tr
383,981
416,458
434,420
1988
386,853
463,457
410,852
0
394,553
471,157
414,652
1989
391,353
391,995
423,837
1,000
393,253
393,895
427,170
1990
385,950
412,154
422,535
5,200
412,450
438,654
434,569
1991
297,831
340,645
381,598
800
298,931
341,745
391,431
1992
257,709
406,457
386,419
300
271,309
420,057
400,152
1993
253,799
378,883
375,328
37,750
321,849
446,933
402,912
1994
395,543
477,215
420,852
0
2,400
410,143
491,815
452,935
a CO, OK, and NM were eliminated from the Official Survey Area in 1985 by the CF CMU.
b The 2018 revision to the Management Plan added OK and WY to counts for determination of the percentage of cranes in the Central Platte River Valley.
c Survey not conducted due to COVID-19 pandemic and associated travel restrictions.
30
Table 1 (continued)
Year
CPRV
Ocular
Cruise
Transect
CPRV
Ocular
Transect
CPRV
Photo
Corrected
Ocular
Transect
Annual
CPRV
Photo
Corrected
Ocular
Transect
3-Year AVG
Other
KS
CO
a
OK
a,b
NM
a
WY
b
Total
Ocular
Cruise
Transect
Total
Ocular
Transect
Total Photo
Corrected
Ocular
Transect
Annual
Total Photo
Corrected
Ocular
Transect
3-Year AVG
1995
273,376
326,181
394,093
0
6,700
303,776
356,581
431,776
1996
318,514
519,984
441,127
0
3,900
326,114
527,584
458,660
1997
350,932
534,630
460,265
100
367,232
550,930
478,365
1998
337,203
530,848
528,487
100
350,903
544,548
541,021
1999
219,794
284,858
450,112
100,000
323,294
388,358
494,612
2000
484,585
490,118
435,275
26,100
528,085
533,618
488,841
2001
387,336
413,498
396,158
42,300
443,636
469,798
463,925
2002
309,029
315,044
406,220
15,100
5,800
342,429
348,444
450,620
2003
300,918
348,023
358,855
4,100
333,618
380,723
399,655
2004
365,370
426,534
363,200
1,200
100
386,470
447,634
392,267
2005
412,285
491,915
422,157
2,900
2,600
450,985
530,615
452,991
2006
178,564
216,810
378,420
2,100
256,164
294,410
424,220
2007
307,094
384,118
364,281
3,600
336,994
414,018
413,014
2008
474,051
545,884
382,271
1,100
499,651
571,484
426,637
2009
457,436
565,257
498,420
tr
498,136
605,957
530,486
2010
455,104
691,534
600,892
1,300
502,004
738,434
638,625
2011
347,501
482,797
579,863
3,500
4,700
384,101
519,397
621,263
2012
253,783
339,642
504,658
tr
270,883
356,742
538,191
2013
745,854
867,061
563,167
279
1,800
771,953
893,160
589,766
2014
402,228
617,903
608,202
5,996
239
440,148
655,823
635,242
a CO, OK, and NM were eliminated from the Official Survey Area in 1985 by the CF CMU.
b The 2018 revision to the Management Plan added OK and WY to counts for determination of the percentage of cranes in the Central Platte River Valley.
c Survey not conducted due to COVID-19 pandemic and associated travel restrictions.
31
Table 1 (continued)
Year
CPRV
Ocular
Cruise
Transect
CPRV
Ocular
Transect
CPRV
Photo
Corrected
Ocular
Transect
Annual
CPRV
Photo
Corrected
Ocular
Transect
3-Year AVG
Other
NE
KS
TX
CO
a
OK
a,b
NM
a
WY
b
Total
Ocular
Cruise
Transect
Total
Ocular
Transect
Total Photo
Corrected
Ocular
Transect
Annual
Total Photo
Corrected
Ocular
Transect
3-Year AVG
2015
326,053
386,471
623,812
24,545
4,479
37,121
2,195
392,198
452,616
667,200
2016
272,250
405,716
470,030
11,218
261
16,500
175
300,229
433,695
514,045
2017
436,671
568,369
453,519
18,674
180
9,193
16
464,718
596,416
494,242
2018
b
516,397
1,005,612
659,899
12,137
1,058
23,906
932
3,475
557,905
1,047,120
692,410
2019
b
633,839
945,996
839,992
16,818
2,423
39,460
777
4,140
697,457
1,009,614
884,383
2020
c
924,289
2021
b
487,418
782,462
911,357
9,394
1,422
38,123
0
4,512
536,339
835,851
964,195
2022
b
464,933
685,476
804,645
23,911
727
13,869
115
3,650
503,407
730,472
858,646
2023
686,716
1,259,199
909,046
6,661
662
11,872
68
6,100
705,895
1,284,563
950,295
a CO, OK, and NM were eliminated from the Official Survey Area in 1985 by the CF CMU.
b The 2018 revision to the Management Plan added OK and WY to counts for determination of the percentage of cranes in the Central Platte River Valley.
c Survey not conducted due to COVID-19 pandemic and associated travel restrictions.
32
Table 2A. Federal Mid-Continent sandhill crane permits issued in the Central Flyway (CF) and Minnesota.
Year
CO
KS
MT
NM
ND
OK
SD
TX
WY
CF
Total
MN
1975
401
158
1,225
4,172
171
198
5,482
56
11,863
1976
341
117
1,195
4,137
265
200
5,060
37
11,352
1977
374
82
1,452
6,294
519
134
4,897
48
13,800
1978
343
209
956
5,798
620
98
5,198
52
13,274
1979
528
159
1,288
4,949
470
63
5,098
43
12,598
1980
437
118
1,082
5,754
510
240
5,239
33
13,413
1981
397
53
1,022
5,796
466
197
5,297
30
13,258
1982
528
147
962
4,714
750
579
4,650
40
12,370
1983
575
175
706
8,033
909
528
7,317
63
18,306
1984
538
113
721
7,436
1,187
544
6,838
43
17,420
1985
555
143
710
6,802
1,102
656
7,417
59
17,444
1986
617
99
595
8,926
1,073
705
7,258
25
19,298
1987
610
128
502
8,778
1,213
517
6,289
30
18,067
1988
512
162
480
6,214
1,472
437
7,053
38
16,368
1989
434
172
430
6,128
1,717
524
8,066
25
17,496
1990
389
143
533
7,268
1,725
646
11,994
22
22,720
1991
501
238
602
3,353
1,618
668
11,142
25
18,147
1992
498
303
582
3,760
1,397
721
9,848
18
17,127
1993
411
575
336
541
4,572
1,277
708
10,407
37
18,864
1994
427
567
320
547
4,790
1,561
636
10,515
49
19,412
1995
571
711
351
564
5,242
1,323
650
10,755
42
20,209
1996
612
837
369
499
5,570
1,391
677
11,334
41
21,330
1997
572
997
325
454
4,934
1,393
757
37,365
b
46
46,845
1998
4,937
b
1,088
270
449
6,082
1,385
951
32,523
b
49
42,797
1999
4,847
b
1,235
279
516
6,050
,1,438
810
33,380
b
52
48,607
2000
5,169
b
1,084
283
493
7,451
1,333
721
44,719
b
58
61,311
2001
5,869
b
1,374
253
509
8,078
1,315
680
49,410
b
72
67,560
2002
5,644
b
1,279
303
496
8,245
c
1,186
619
37,558
b
54
55,384
2003
a
5,854
b
1,206
273
471
6,030
c
1,000
563
43,199
b
50
58,646
2004
a
5,784
b
1,180
c
308
548
5,788
c
780
c
307
52,161
b
61
66,917
2005
a
5,766
b
805
c
281
494
7,441
c
698
c
490
51,511
b
68
67,554
2006
a
4,792
b
826
c
265
512
d
7,410
c
615
c
445
e
70,968
b
78
85,911
2007
a
4,931
b
598
c
238
480
d
7,442
c
731
c
390
e
101,382
b
58
116,250
2008
a
5,772
b
655
c
272
677
d
6,501
c
736
c
398
e
122,553
b
73
137,637
2009
a
4,038
b
540
c
139
862
d
7,774
c
1,029
c
693
e
11,332
b
62
26,469
2010
a
4,280
b
508
c
283
701
d
8,375
c
1,055
c
410
e
12,560
b
86
28,258
1,954
2011
a
783
b
801
c
311
575
d
8,024
c
1,104
c
356
e
13,905
b
86
25,945
1,342
2012
a
801
b
571
c
186
859
d
8,519
c
451
c
343
e
14,083
b
102
25,915
1,032
2013
a
856
b
735
c
288
404
d
9,085
c
2,278
c
421
e
18,369
b
106
32,542
1,086
2014
a
848
b
787
c
356
368
d
4,692
c
660
c
390
e
20,105
b
433
28,639
1,216
2015
a
787
b
1,040
c
404
365
d
4,543
c
510
c
---
f
22,033
b
454
30,136
1,199
2016
a
841
b
1,055
c
376
416
d
3,956
c
559
c
171
e
23,962
b
569
31,905
1,139
2017
a
913
b
1,075
c
604
534
d
4,006
c
714
c
224
e
26,312
b
646
35,028
1,125
2018
a
954
b
1,218
c
676
2,413
d
4,102
c
642
c
237
e
29,668
b
392
40,302
1,091
2019
a
1,019
b
1,456
c
1,013
b
2,818
d
3,839
c
---
g
242
e
32,841
b
714
43,942
1,073
2020
a
1,107
b
1,970
c
1,005
b
2,763
d
5,168
c
11,513
c
210
e
38,832
b
619
63,187
1,288
2021
a
1,170
b
2,343
c
1,385
b
2,362
d
4,440
c
11,904
c
289
e
45,013
b
621
69,527
1,479
2022
a
9,194
b
2,270
c
562
b
3,471
d
5,544
c
12,536
c
250
e
52,568
b
1,182
87,577
1,424
a Preliminary
b Harvest Information Program (HIP) or a point-of-sale electronic record (without cost) used to identify crane hunters in lieu of a
special sandhill crane hunting permit
c States began charging a fee for crane hunting permits which reduces the number of permits issued to hunters that only hunt
cranes.
d NM uses a combination of electronic and paper permits.
e SD uses a special question in their HIP questionnaire to identify sandhill crane hunters; TX hunters can only obtain crane permits
in selected locations.
f All hunters put in stratum "did not hunt" or "no" in state HIP sample frame, so no estimate is available.
g Hunters name and address data not supplied, so no estimate is available.
33
Table 2B. Decadal and long-term annual averages of federal Mid-Continent sandhill crane permits issued
in the Central Flyway (CF) and Minnesota.
Years
CO
KS
MT
NM
ND
OK
SD
TX
WY
CF Total
MN
1975-79
397
145
1,223
5,070
409
139
5,147
47
12,577
1980-89
520
131
721
6,858
1,040
493
6,542
39
16,344
1990-99
981
859
293
529
5,162
1,451
722
17,926
38
27,606
2000-09
5,362
955
262
554
7,216
942
531
58,479
63
74,364
2010-19
1,208
925
450
945
5,914
886
310
21,384
359
32,261
1,226
2020-22
3,824
2,194
984
2,865
5,051
11,984
250
45,471
807
73,430
1,397
1975-2022
1,983
1,046
313
879
6,083
1,709
462
25,114
159
37,269
1,265
34
Table 3A. Estimated active Mid-Continent sandhill crane hunters in the Central Flyway (CF) and
Minnesota. Active hunters are those permittees reporting hunting cranes 1 or more times.
Year
CO
KS
MT
NM
ND
OK
SD
TX
WY
CF
Total
MN
1975
226
69
806
2,896
80
117
2,733
22
6,949
1976
203
68
752
1,328
148
80
2,497
16
5,092
1977
189
40
921
4,126
339
77
2,329
27
8,048
1978
190
86
836
3,776
334
50
2,390
21
7,683
1979
275
61
745
3,225
307
29
2,356
13
7,011
1980
216
50
625
3,387
275
160
2,439
12
7,164
1981
216
23
598
3,315
269
103
2,543
14
7,081
1982
138
56
386
2,429
342
260
1,553
8
5,172
1983
211
64
253
3,551
384
225
2,435
20
7,143
1984
206
51
301
3,189
467
208
2,380
19
6,821
1985
187
37
216
2,383
372
168
2,613
12
5,988
1986
106
17
178
3,095
299
149
1,991
5
5,840
1987
113
29
133
2,529
358
120
1,942
5
5,229
1988
117
48
171
1,779
531
78
2,497
11
5,232
1989
74
52
152
2,018
492
153
2,805
6
5,752
1990
101
33
180
2,614
395
172
4,130
6
7,631
1991
153
69
220
1,674
370
139
3,231
3
5,859
1992
96
95
182
1,776
330
153
2,655
7
5,294
1993
87
294
97
218
2,223
357
140
3,602
5
7,023
1994
93
293
79
211
2,497
456
151
3,350
11
7,141
1995
154
393
118
211
2,408
331
143
3,707
6
7,471
1996
91
382
82
166
2,744
355
169
3,356
9
7,354
1997
67
452
68
124
2,386
264
178
4,515
10
8,064
1998
96
480
43
155
2,785
345
237
4,022
10
8,173
1999
133
533
60
204
2,444
375
173
2,699
8
6,629
2000
192
430
64
160
2,481
223
209
3,180
11
6,950
2001
202
555
72
173
2,934
391
145
3,554
13
8,039
2002
175
517
85
166
2,407
237
144
4,037
15
7,783
2003
a
236
495
60
244
2,271
64
114
4,821
10
8,315
2004
a
315
539
93
252
2,491
265
79
5,121
16
9,171
2005
a
280
274
90
233
3,370
259
165
5,383
24
10,078
2006
a
144
445
71
245
3,272
243
144
5,531
25
10,120
2007
a
158
255
82
241
3,145
166
57
5,685
19
9,808
2008
a
191
283
84
239
2,815
255
64
6,338
24
10,293
2009
a
159
213
50
286
3,546
371
63
3,179
67
7,934
2010
a
302
182
93
192
3,474
332
52
4,187
29
8,843
964
2011
a
138
449
95
206
3,733
418
44
2,712
41
7,836
643
2012
a
139
214
59
270
3,332
160
54
2,972
39
7,239
410
2013
a
118
235
94
276
3,326
638
91
5,473
35
10,286
485
2014
a
89
151
88
252
1,743
231
56
5,145
70
7,825
401
2015
a
126
334
115
263
1,430
158
---
b
3,241
78
5,745
424
2016
a
144
332
113
310
1,504
219
39
6,746
96
9,503
471
2017
a
221
710
98
360
1,562
246
71
7,066
305
10,639
397
2018
a
178
457
175
416
1,626
258
73
8,807
94
12,084
383
2019
a
174
554
152
549
1,124
---
c
41
10,072
138
12,804
333
2020
a
216
735
229
505
1,752
3,722
52
19,999
177
27,387
480
2021
a
251
818
358
498
1,633
3,116
71
14,240
116
21,101
632
2022
a
768
504
240
579
2,272
1,947
50
16,822
347
23,529
898
a
Preliminary
b
All hunters put in stratum "did not hunt" or "no" in state HIP sample frame, so no estimate is available.
c
Hunter name and address data not supplied, so no estimate is available.
35
Table 3B. Decadal and long-term annual averages of active Mid-Continent sandhill crane hunters in the
Central Flyway (CF) and Minnesota.
Years
CO
KS
MT
NM
ND
OK
SD
TX
WY
CF Total
MN
1975-79
217
65
812
3,070
242
71
2,461
20
6,957
1980-89
158
43
301
2,768
379
162
2,320
11
6,142
1990-99
107
404
74
187
2,355
358
166
3,527
8
7,064
2000-09
205
401
75
224
2,873
247
118
4,683
22
8,849
2010-19
163
362
108
309
2,285
296
58
5,642
93
9,280
491
2020-22
412
686
276
527
1,886
2,928
58
17,020
213
24,006
670
1975-22
180
417
87
330
2,580
479
117
4,689
43
8,753
532
36
Table 4. Season dates (month/day) for the hunting of Mid-continent sandhill cranes in the Central Flyway states and Minnesota.
Year
CO
KS
Central
Zone¹
KS
West
Zone²
MT¹
MT²
NM
ND
Area 1
ND
Area 2
OK
SD
TX
Area A
TX
Area B
TX
Area C
WY
MN
1960
-
-
-
-
-
01/01-01/30
-
-
-
-
-
-
-
-
-
1961
-
-
-
-
-
11/04-12/03
-
-
-
-
11/04-12/03
-
-
-
-
1962
-
-
-
-
-
11/03-12/02
-
-
-
-
11/03-12/02
-
-
-
-
1963
-
-
-
-
-
11/02-12/01
-
-
-
-
11/02-12/01
-
-
-
-
1964
-
-
-
-
-
10/31-11/29
-
-
-
-
10/31-11/29
-
-
-
-
1965
-
-
-
-
-
10/30-11/28
-
-
-
-
10/30-11/28
-
-
-
-
1966
-
-
-
-
-
10/29-11/27
-
-
-
-
10/29-11/27
-
-
-
-
1967
10/01-10/30
-
-
-
-
11/04-01/02
-
-
-
-
11/04-01/02
-
-
-
-
1968
10/01-10/30
-
-
-
-
11/02-12/28
11/09-12/08
-
12/14-01/02
11/09-12/08
11/02-12/28
12/14-01/02
-
-
-
1969
10/04-11/02
-
-
-
-
11/01-12/28
11/08-12/07
-
12/13-01/11
11/08-12/07
11/01-12/28
12/13-01/11
-
-
-
1970
10/03-11/01
-
-
-
-
10/31-01/10
11/14-12/13
-
12/05-01/10
11/14-12/13
10/31-01/10
12/05-01/10
-
-
-
1971
10/02-11/07
-
-
-
-
10/30-01/30
11/13-12/02
-
12/04-01/30
11/13-12/02
10/30-01/30
12/04-01/30
-
-
-
1972
10/01-11/05
-
-
10/01-11/06
-
11/03-01/31
11/11-12/10
-
12/02-01/28
11/11-12/10
10/28-01/28
12/02-01/28
-
10/07-11/05
-
1973
10/01-11/05
-
-
09/29-11/04
-
10/27-01/27
11/10-12/09
-
12/01-01/27
11/10-12/09
10/27-01/27
12/01-01/27
-
10/13-11/11
-
1974
10/01-11/05
-
-
09/28-11/03
-
10/26-01/26
11/09-12/08
-
11/30-01/26
11/09-12/08
10/26-01/26
11/30-01/26
-
10/12-11/10
-
1975
10/04-11/08
-
-
10/04-11/09
-
10/25-01/25
11/08-12/07
-
11/29-01/25
11/08-12/07
10/25-01/25
11/29-01/25
-
10/11-11/09
-
1976
10/02-11/06
-
-
10/02-11/07
-
10/30-01/30
11/06-12/05
-
11/27-01/23
11/06-12/05
10/30-01/30
12/04-01/30
-
10/09-11/07
-
1977
10/01-11/06
-
-
10/01-11/06
-
10/29-01/29
09/07-09/11
-
11/26-01/22
09/07-09/11
11/01-01/31
12/05-01/31
-
10/08-11/06
-
1978
09/30-11/05
-
-
09/30-11/05
-
10/28-01/28
09/07-09/11
-
11/25-01/21
09/07-09/11
10/31-01/31
12/05-01/31
-
10/07-11/05
-
1979
10/13-11/18
-
-
09/29-11/04
-
10/27-01/27
09/07-09/11
-
11/24-01/20
09/07-09/11
10/30-01/30
12/04-01/30
-
10/13-11/18
-
1980
10/11-11/16
-
-
10/04-11/09
-
10/30-01/31
09/06-09/14
09/06-09/10
11/22-01/18
09/20-09/28
10/31-01/31
12/05-01/31
-
10/11-11/16
-
1981
10/10-11/15
-
-
10/03-11/08
-
10/31-01/31
09/05-09/20
09/05-09/13
11/22-01/18
09/20-09/28
10/31-01/31
12/05-01/31
-
10/03-11/08
-
1982
10/02-11/28
-
-
10/02-11/28
-
10/31-01/31
09/04-09/19
09/04-09/12
10/23-01/23
10/02-11/11
10/30-01/30
12/04-01/30
-
09/25-11/21
-
1983
10/01-11/27
-
-
11/01-11/27
11/01-11/27
10/29-01/28
09/10-11/06
09/10-09/30
10/22-01/22
10/01-11/06
11/12-02/12
12/03-02/12
01/14-02/12
09/24-11/20
-
1984
09/29-11/25
-
-
09/29-11/25
11/01-11/25
10/27-01/27
09/08-11/04
09/08-09/28
10/13-01/13
09/29-11/04
11/10-02/10
12/01-02/10
01/12-02/10
09/22-11/18
-
1985
09/28-11/24
-
-
09/28-11/24
11/01-11/24
10/26-01/26
09/07-11/03
09/07-09/27
10/12-01/12
09/28-11/03
11/09-02/09
11/30-02/09
01/11-02/09
09/21-11/17
-
1986
10/04-11/30
-
-
10/04-11/30
11/01-11/30
10/25-01/25
09/06-11/02
09/06-10/03
10/11-01/11
09/28-11/02
11/08-02/08
11/29-02/08
01/03-02/08
09/20-11/16
-
1987
10/03-11/29
-
-
10/03-11/29
10/03-11/29
10/24-01/24
09/05-11/01
09/05-10/02
10/10-01/17
09/26-11/01
11/14-02/14
11/28-02/07
01/02-02/07
09/19-11/15
-
1988
10/01-11/27
-
-
10/01-11/27
10/01-11/27
10/22-01/22
09/10-11/06
09/10-09/30
10/22-01/22
09/24-10/30
11/12-02/12
11/26-02/05
01/07-02/12
09/17-11/13
-
1989
09/30-11/26
-
-
09/30-11/26
09/30-11/26
10/21-01/21
09/09-11/05
09/09-09/29
10/21-01/21
09/30-11/05
11/11-02/11
12/02-02/11
01/06-02/11
09/16-11/12
-
1990
09/29-11/25
-
-
09/29-11/25
09/29-11/25
10/20-01/20
09/08-11/04
09/08-10/14
10/20-01/20
09/29-11/04
11/10-02/10
12/01-02/10
01/05-02/10
09/15-11/11
-
1991
09/28-11/24
-
-
09/28-11/24
09/28-11/24
10/19-01/19
09/07-11/03
09/07-10/13
10/19-01/19
09/28-11/03
11/09-02/09
12/07-02/09
01/04-02/09
09/15-11/11
-
1992
10/03-11/29
-
-
09/26-11/22
09/26-11/22
10/17-01/17
09/05-11/01
09/05-10/11
10/17-01/17
09/26-11/01
11/14-02/14
12/05-02/14
01/02-02/07
09/15-11/11
-
1993
10/02-11/28
11/06-01/02
-
09/25-11/21
09/25-11/21
10/16-01/16
09/11-11/07
09/11-11/07
10/16-01/16
09/25-10/31
11/13-02/13
12/04-02/13
01/08-02/13
09/15-11/11
-
1994
10/01-11/27
11/05-01/01
-
09/24-11/20
09/24-11/20
10/15-01/15
09/10-11/06
09/10-11/06
10/15-01/15
09/24-10/30
11/12-02/12
12/03-02/12
01/07-02/12
09/15-11/11
-
1995
09/30-11/26
11/04-12/31
-
09/23-11/19
09/23-11/19
10/31-01/31
09/09-11/05
09/09-11/05
10/22-01/28
09/23-11/19
11/11-02/11
12/02-02/11
01/06-02/11
09/14-11/10
-
KS
1
Central Zone: East of the West Zone Boundary to the line of OK/KS state line headed north on I-35 to I-135, I-70 to US 81 to the NE state line.
KS
2
West Zone: KS/NE State Line south along US 283, KS road 24, KS -18 to US 183 to KS - 1 to the KS/OK state line.
MT¹ Central Flyway portion of MT, except that area south of I-90 and west of the Bighorn River and Sheridan Co.
MT² Sheridan County, MT.
37
Table 4 (continued)
Year
CO
KS
Central
Zone¹
KS
West
Zone²
MT¹
MT²
NM
ND
Area 1
ND
Area 2
OK
SD
TX
Area A
TX
Area B
TX
Area C
WY
MN
1996
10/05-12/01
11/02-12/29
-
09/28-11/24
09/28-11/24
10/31-01/31
09/07-11/03
09/07-11/03
10/26-01/26
09/28-11/24
11/09-02/09
11/30-02/09
01/04-02/09
09/14-11/10
-
1997
10/04-11/30
11/01-12/28
-
10/04-11/30
10/04-11/30
10/31-01/31
09/06-11/02
09/06-11/02
10/25-01/25
09/27-11/23
11/08-02/08
11/29-02/08
01/03-02/08
09/13-11/09
-
1998
10/03-11/29
11/07-01/03
-
10/03-11/29
09/12-09/20
10/31-01/31
09/05-11/01
09/05-11/01
10/24-01/24
09/26-11/22
11/07-02/07
11/28-02/07
01/02-02/07
09/12-11/08
-
1999
10/02-11/28
11/06-01/02
-
10/02-11/28
09/11-09/19
10/30-01/30
09/11-11/07
09/11-11/07
10/30-01/30
09/25-11/21
11/13-02/13
12/04-02/13
01/08-02/13
09/11-11/07
-
2000
10/07-12/03
11/04-12/31
-
09/30-11/26
09/09-09/17
10/31-01/31
09/16-11/12
09/16-11/12
11/04-02/04
09/23-11/19
11/11-02/11
12/02-02/11
12/30-02/04
09/09-11/05
-
2001
10/07-12/03
11/03-12/30
-
09/29-11/25
09/08-09/16
10/31-01/31
09/15-11/11
09/15-10/21
11/03-02/03
09/22-11/18
11/10-02/10
12/01-02/10
12/29-01/20
09/15-11/11
-
2002
10/05-12/01
11/02-12/29
-
09/28-11/24
09/07-09/15
10/31-01/31
09/21-11/17
09/21-10/27
11/09-02/09
09/21-11/17
11/09-02/09
11/30-02/09
12/21-01/19
09/14-11/10
-
2003
10/04-11/30
11/01-12/28
-
09/27-11/23
09/06-09/14
10/31-01/31
09/20-11/16
09/20-10/26
10/25-01/25
09/27-11/23
11/01-02/01
11/22-02/01
12/20-01/18
09/13-11/09
-
2004
10/02-11/28
11/06-01/02
-
09/25-11/21
09/11-09/19
10/31-01/31
09/18-11/14
09/18-10/24
10/30-01/30
09/25-11/21
11/06-02/01
11/27-02/01
12/18-01/16
09/18-11/14
-
2005
10/01-11/27
11/09-01/05
-
09/24-11/20
09/10-09/18
10/31-01/31
09/17-11/13
09/17-10/23
10/29-01/29
09/24-11/20
11/05-02/05
11/26-02/05
12/24-01/29
09/17-11/13
-
2006
09/30-11/26
11/08-01/04
-
09/23-11/19
09/09-09/17
10/31-01/31
09/16-11/12
09/16-10/22
10/28-01/28
09/23-11/19
11/04-02/04
11/24-02/04
12/23-01/28
09/16-11/12
-
2007
10/02-12/02
11/07-01/03
-
09/22-11/18
09/08-09/16
10/31-01/31
09/15-11/11
09/15-10/21
10/27-01/27
09/22-11/18
11/04-02/04
11/24-02/04
12/23-01/28
09/15-11/11
-
2008
10/04-11/30
11/05-01/01
-
09/27-11/23
09/06-09/21
10/31-01/31
09/20-11/16
09/20-10/26
10/25-01/25
09/27-11/23
11/08-02/08
11/28-02/08
12/20-01/25
09/13-11/09
-
2009
10/03-11/29
11/11-01/07
-
09/26-11/22
09/05-09/20
10/31-01/31
09/19-11/15
09/19-10/25
10/24-01/24
09/26-11/22
11/07-02/07
11/27-02/07
12/19-01/24
09/19-11/15
-
2010
10/02-11/28
11/10-01/06
-
09/25-11/21
09/11-09/26
10/31-01/31
09/18-11/14
09/18-10/24
10/23-01/23
09/25-11/21
11/06-02/06
11/26-02/06
12/18-01/23
09/18-11/14
09/04-10/10
2011
10/01-11/27
11/09-01/05
-
09/24-11/20
09/10-09/25
10/31-01/31
09/17-11/13
09/17-10/23
10/22-01/22
09/24-11/20
11/05-02/05
11/25-02/05
12/24-01/29
09/17-11/13
09/03-10/09
2012
09/29-11/25
11/07-01/03
-
09/29-11/25
09/8-09/30
10/31-01/31
09/15-11/11
09/15-10/21
10/20-01/20
09/22-11/18
11/03-02/03
11/23-02/03
12/22-01/27
09/15-11/11
09/15-10/21
2013
10/05-12/01
11/06-01/02
-
09/28-11/24
09/07-09/29
10/31-01/31
09/14-11/10
09/14-11/10
10/19-01/19
09/28-11/24
11/02/-02/02
11/22-02/02
12/21-01/26
09/14-11/10
09/14-10/20
2014
10/04-11/30
11/05-01/01
-
10/04-11/30
09/13-10/05
10/31-01/31
09/14-11/10
09/14-11/10
10/18-01/18
09/27-11/23
11/01/-02/01
11/21-02/01
12/20-01/25
09/13-11/09
09/13-10/19
2015
10/03-11/29
11/11-01/07
-
10/03-11/29
09/12-10/04
10/31-01/31
09/19-11/15
09/19-11/15
10/24-01/24
09/26-11/22
10/31-01/31
11/20-01/31
12/19-01/24
09/19-11/15
09/12-10/18
2016
10/01-11/27
11/09-01/05
-
10/01-11/27
09/10-10/02
10/29-01/29
09/17-11/13
09/17-11/13
10/22-01/22
09/24-11/20
10/29-01/29
11/18-01/29
12/17-01/22
09/17-11/13
09/10-10/16
2017
09/30-11/26
11/08-01/04
-
09/30-11/26
09/09-10/08
10/28-01/28
09/16-11/12
09/16-11/12
10/21-01/21
09/23-11/19
10/28-01/28
11/24-01/28
12/16-01/21
09/16-11/12
09/16-10/22
2018
09/29-11/25
11/07-01/03
-
09/29-11/25
09/01-10/28
10/27/01/27
09/15-11/11
09/15-11/11
10/20-01/20
09/22-11/18
10/27-01/27
11/23-01/27
12/15-01/20
09/15-11/11
09/15-10/21
2019
10/05-12/01
11/06-01/02
-
09/28-11/24
09/01-10/27
10/26-01/26
09/14-11/10
09/14-11/10
10/26-01/26
09/28-11/24
10/26-01/26
11/22-01/26
12/14-01/19
09/14-11/10
09/14-10/20
2020
10/03-11/29
11/11-01/07
10/17-12/13
10/03-11/29
09/01-10/30
10/24-01/24
09/19-11/15
09/19-11/15
10/24-01/24
09/26-11/22
10/31-01/31
11/27-01/31
12/19-01/24
09/12-11/08
09/19-10/25
2021
10/02-11/28
11/10-01/06
10/16-12/12
10/02-11/28
09/01-10/30
10/30-01/30
09/18-11/14
09/18-11/14
10/23-01/23
09/25-11/21
10/30-01/30
11/26-01/30
12/18/01/23
09/11-11/07
09/18-10/24
2022
10/01-11/27
11/09-01/05
10/15-12/11
10/01-11/27
09/01-10/30
10/29-01/29
09/17-11/13
09/17-11/13
10/22-01/22
09/24-11/20
10/29-01/29
11/25-01/29
12/17-01/22
09/10-11/06
09/17-10/23
KS
1
Central Zone: East of the West Zone Boundary to the line of OK/KS state line headed north on I-35 to I-135, I-70 to US 81 to the NE state line.
KS
2
West Zone: KS/NE State Line south along US 283, KS road 24, KS -18 to US 183 to KS - 1 to the KS/OK state line.
MT¹ Central Flyway portion of MT, except that area south of I-90 and west of the Bighorn River and Sheridan Co.
MT² Sheridan County, MT.
38
Table 5A. Estimated retrieved harvests of Mid-Continent sandhill cranes in the Central Flyway states, and states outside the Central Flyway (AZ,
NM, AK, MN)
.
Year
CO
KS
MT
NM
ND
OK
SD
TX
WY
Central
Flyway
AZ
d
NM
d
AK
b,c
MN
e
Outside
Total
U.S.
Total
1975
91
16
911
2,122
142
86
6,123
6
9,497
1,094
1,094
10,591
1976
106
29
858
52
200
12
6,122
14
7,393
637
637
8,030
1977
39
18
1,456
4,078
410
47
6,094
9
12,151
471
471
12,622
1978
106
36
1,089
2,777
389
19
5,720
10
10,146
239
239
10,385
1979
129
14
1,170
2,733
397
19
5,917
0
10,379
517
517
10,896
1980
68
16
1,019
2,245
363
130
6,305
6
10,152
809
809
10,961
1981
92
11
907
2,395
397
78
6,245
9
10,134
20
383
403
10,537
1982
49
21
335
2,469
535
212
4,295
0
7,916
62
1,160
1,222
9,138
1983
70
28
354
6,471
373
177
5,471
15
12,959
17
1,540
1,557
14,516
1984
85
15
414
4,367
433
139
5,811
7
11,271
23
1,986
2,009
13,280
1985
82
7
334
4,650
416
101
7,184
2
12,776
48
1,197
1,245
14,021
1986
33
1
250
6,563
392
99
5,149
0
12,487
108
184
539
831
13,318
1987
86
15
159
5,334
957
99
6,117
3
12,770
127
318
836
1,281
14,051
1988
68
18
372
3,815
1,061
100
7,330
8
12,772
172
127
1,241
1,540
14,312
1989
25
33
319
4,656
1,003
194
7,400
9
13,639
126
138
545
809
14,448
1990
87
44
377
6,804
698
165
9,865
1
18,041
114
259
918
1,291
19,332
1991
224
31
593
4,580
604
128
6,916
3
13,079
172
235
677
1,084
14,163
1992
84
103
505
4,654
478
141
6,455
13
12,433
139
54
640
833
13,266
1993
112
602
95
506
6,985
826
110
8,769
0
18,005
113
178
201
492
18,497
1994
143
767
56
357
6,235
1,167
239
7,233
4
16,201
86
153
648
887
17,088
1995
208
990
156
673
7,017
1,091
170
10,322
1
20,628
124
111
812
1,047
21,675
1996
91
933
58
332
6,639
1,066
166
7,816
10
17,111
114
78
1,205
1,397
18,508
1997
168
1,167
45
248
6,545
600
189
10,800
4
19,766
171
45
870
1,086
20,852
1998
64
1,362
17
258
7,967
645
454
9,054
10
19,831
114
55
1,042
1,211
21,042
1999
56
1,275
29
321
5,748
879
184
8,469
8
16,969
92
101
NA*
193
17,162
2000
363
590
15
311
5,081
552
374
8,208
10
15,504
166
100
985
1,251
16,755
2001
257
1,033
43
297
5,173
713
478
6,999
7
15,000
154
106
936
1,196
16,196
2002
294
1,067
23
342
2,852
490
160
7,837
22
13,087
197
92
844
1,133
14,220
a
Preliminary
b
A proportion of the Alaskan harvest is composed of lesser sandhill cranes from the Pacific Coast Population
c
Harvest data are from state harvest surveys for only the MCP portion of the state, except in 1977-81, 1986, 1991, and 1998-99 where
federal MQS state totals are prorated by the long-term percent MC cranes; data from 2000 forward are MC portion from HIP.
d
The MC harvest for AZ and NM represents MC sandhill cranes that were harvested in RMP areas and are not represented in the CF MC sandhill Crane Federal Harvest Survey
e
Minnesota initiated a hunt in the NW portion of state.
g
HIP sample frame from state was incomplete, so no estimate was calculated.
* No estimate is available.
39
Table 5A (continued)
Year
CO
KS
MT
NM
ND
OK
SD
TX
WY
Central
Flyway
AZ
d
NM
d
AK
b,c
MN
e
Outside
Total
U.S.
Total
2003
a
230
942
49
617
4,564
200
166
11,560
7
18,335
155
162
331
648
18,983
2004
a
92
856
54
350
3,967
441
67
8,715
4
14,546
192
167
435
794
15,340
2005
a
265
471
65
578
3,721
511
190
12,446
16
18,263
227
175
388
790
19,053
2006
a
96
1,341
12
682
3,906
538
202
10,834
20
17,631
201
245
314
760
18,391
2007
a
149
516
51
427
4,501
272
163
12,511
20
18,610
268
331
596
1,195
19,805
2008
a
32
453
73
483
4,179
493
83
17,169
24
22,989
138
329
1,249
1,716
24,705
2009
a
58
447
34
584
4,436
737
96
8,882
8
15,282
305
332
245
882
16,164
2010
a
115
293
95
432
4,752
940
91
12,069
25
18,812
253
421
1,204
830
2,708
21,520
2011
a
68
908
51
297
3,733
808
64
8,493
20
14,442
151
367
335
765
1,618
16,060
2012
a
77
437
30
388
3,019
401
185
10,309
41
14,887
300
341
1,360
407
2,408
17,295
2013
a
47
771
77
326
4,137
1,085
109
14,991
41
21,584
138
161
930
378
1,607
23,191
2014
a
41
176
114
269
2,924
390
85
11,740
37
15,776
151
123
1,123
247
1,644
17,420
2015
a
98
1,005
91
267
2,133
302
---
f
8,283
28
12,207
311
132
---
g
212
655
12,862
2016
a
102
873
111
660
2,507
538
183
18,196
83
23,253
292
404
1,036
287
2,019
25,272
2017
a
280
1,440
85
641
3,466
559
165
19,559
263
26,458
435
399
793
196
1,823
28,281
2018
a
102
1,127
73
701
3,424
718
119
22,526
33
28,823
587
284
705
129
1,705
30,528
2019
a
131
1,160
82
1,326
1,764
---
43
29,607
82
34,195
187
264
659
179
1,289
35,484
2020
a
229
1,718
95
1,520
2,998
8,974
146
55,871
182
71,733
651
671
1,409
472
3,203
74,936
2021
a
212
2,012
339
1,336
4,470
7,791
109
43,217
79
59,565
722
345
1,577
563
3,207
62,772
2022
a
371
631
237
1,721
3,465
3,164
75
45,419
233
55,316
736
269
794
863
2,662
57,978
a
Preliminary
b
A proportion of the Alaskan harvest is composed of lesser sandhill cranes from the Pacific Coast Population
c
Harvest data are from state harvest surveys for only the MCP portion of the state, except in 1977-81, 1986, 1991, and 1998-99 where
federal MQS state totals are prorated by the long-term percent MC cranes; data from 2000 forward are MC portion from HIP.
d
The MC harvest for AZ and NM represents MC sandhill cranes that were harvested in RMP areas and are not represented in the CF MC sandhill Crane Federal Harvest Survey
e
Minnesota initiated a hunt in the NW portion of state.
f
All hunters put in stratum "did not hunt" or "no" in state HIP sample frame, so no estimate is available.
g
HIP sample frame from state was incomplete, so no estimate was calculated.
* No estimate is available.
40
Table 5B. Decadal and long-term annual averages of harvest of active Mid-Continent sandhill cranes in the Central Flyway states (CO, KS, MT, NM,
ND, OK, SD, TX, and WY), states outside the Central Flyway (AZ, NM, AK, MN).
Years
CO
KS
MT
NM
ND
OK
SD
TX
WY
Central
Flyway
Total
AZ
d
NM
d
AK
b,c
MN
e
Outside
Total
U.S.
Total
1975-79
94
23
1,097
2,352
308
37
5,995
8
9,913
592
592
10,505
1980-89
66
17
446
4,297
593
133
6,131
6
11,688
78
192
1,024
1,171
12,858
1990-99
124
1,014
63
417
6,317
805
195
8,570
5
17,206
124
127
779
952
18,159
2000-09
184
772
42
467
4,238
495
198
10,516
14
16,925
200
204
632
1,037
17,961
2010-19
106
819
81
531
3,186
638
116
15,577
65
21,044
281
290
905
389
1,748
22,791
2020-21
271
1,454
224
1,526
3,644
6,643
110
48,169
165
62,492
703
428
1,260
633
3,024
65,516
1975-2021
127
912
59
597
4,231
982
145
12,134
30
18,868
211
223
836
425
1,273
20,141
41
Table 6. Estimated retrieved harvests of Mid-Continent sandhill cranes in Canada.
Year
AB
MB
SK
Total
1971
0
228
2,715
2,943
1972
0
113
2,030
2,143
1973
0
683
3,592
4,275
1974
0
58
6,641
6,699
1975
0
162
5,744
5,906
1976
0
868
1,767
2,635
1977
0
856
510
1,366
1978
0
1,111
421
1,532
1979
0
1,113
765
1,878
1980
0
1,412
5,688
7,100
1981
0
1,317
4,746
6,063
1982
0
1,191
3,794
4,985
1983
0
1,205
5,012
6,217
1984
0
1,210
5,197
6,407
1985
0
1,268
6,805
8,073
1986
0
1,577
8,463
10,040
1987
0
1,812
6,135
7,947
1988
0
1,698
5,394
7,092
1989
0
2,268
4,321
6,589
1990
0
1,848
5,351
7,199
1991
0
1,513
5,420
6,933
1992
0
1,476
5,242
6,718
1993
0
1,086
4,137
5,223
1994
0
1,224
4,925
6,149
1995
0
1,287
7,072
8,359
1996
0
1,519
6,058
7,577
1997
0
1,481
7,075
8,556
1998
0
1,557
8,640
10,197
1999
0
1,504
7,580
9,084
2000
0
1,473
9,082
10,555
2001
0
1,861
8,132
9,993
2002
0
2,043
7,105
9,148
2003
0
2,333
7,810
10,143
2004
0
2,595
8,951
11,546
2005
0
3,239
8,897
12,136
2006
0
3,549
8,366
11,915
2007
0
3,364
8,514
11,878
2008
0
2,862
8,959
11,821
2009
0
2,454
5,758
8,212
2010
0
2,659
8,021
10,680
2011
0
3,271
9,939
13,210
2012
0
3,294
8,746
12,040
2013
0
3,472
9,860
13,332
2014
0
3,629
11,782
15,411
2015
0
3,655
10,804
14,459
2016
0
3,739
10,499
14,238
2017
0
2,905
12,456
15,361
2018
0
3,120
11,210
14,330
2019
0
2,671
13,100
15,771
2020
969
2,065
5,233
7,298
2021
959
2,229
9,850
12,079
2022
a
a
Estimates not available at the time the status report was completed.
42
Table 7A. Annual sport hunting mortality estimates for the Mid-Continent Population of sandhill cranes in
North America.
Year
Central
Flyway
Other Survey
Area Total
Canada
Mexico
b
Unretrieved
No. America
c
Total
1975
9,497
1,094
5,906
1,650
3,615
21,762
1976
7,393
637
2,635
1,067
2,252
13,983
1977
12,151
471
1,366
1,399
2,660
18,047
1978
10,146
239
1,532
1,192
2,453
15,561
1979
10,379
517
1,878
1,277
2,384
16,436
1980
10,152
809
7,100
1,806
3,683
23,550
1981
10,134
403
6,063
1,660
3,405
21,665
1982
7,916
1,222
4,985
1,412
2,924
18,459
1983
12,959
1,557
6,217
2,073
4,189
26,996
1984
11,271
2,009
6,407
1,969
3,967
25,623
1985
12,776
1,245
8,073
2,209
4,165
28,469
1986
12,487
831
10,040
2,336
4,511
30,205
1987
12,770
1,281
7,947
2,200
3,996
28,193
1988
12,772
1,540
7,092
2,140
3,782
27,327
1989
13,639
809
6,589
2,104
3,981
27,122
1990
18,041
1,291
7,199
2,653
4,748
33,932
1991
13,079
1,084
6,933
2,110
3,793
26,999
1992
12,433
833
6,718
1,998
3,369
25,351
1993
18,005
492
5,223
2,372
3,869
29,961
1994
16,201
887
6,149
2,324
3,548
29,109
1995
20,628
1,047
8,359
3,003
4,718
37,755
1996
17,111
1,397
7,577
2,609
4,328
33,021
1997
19,766
1,086
8,556
2,941
4,796
37,144
1998
19,831
1,211
10,197
3,124
5,048
39,411
1999
16,969
193
4
9,084
2,625
4,097
32,968
2000
15,504
1,251
10,555
2,731
4,336
34,377
2001
15,000
1,196
9,993
2,619
4,279
33,086
2002
13,087
1,133
9,148
2,337
3,711
29,415
2003
a
18,335
648
10,143
2,913
4,369
36,407
2004
a
14,546
794
11,546
2,689
4,277
33,851
2005
a
18,263
790
12,136
3,119
5,009
39,317
2006
a
17,631
760
11,915
3,031
5,193
38,530
2007
a
18,610
1,195
11,878
3,168
4,924
39,776
2008
a
22,989
1,716
11,821
3,653
4,956
45,135
2009
a
15,282
882
8,212
2,438
3,990
30,804
2010
a
18,812
2,708
10,680
3,220
5,137
40,557
2011
a
14,442
1,618
13,210
2,927
4,617
36,814
2012
a
14,887
2,408
12,040
2,934
4,936
37,205
2013
a
21,584
1,607
13,332
3,652
4,947
45,123
2014
a
15,776
1,644
15,411
3,283
5,093
41,207
2015
a,e
12,207
655
14,459
2,732
4,500
34,553
2016
a
23,253
2,019
14,238
3,951
5,062
48,523
2017
a
26,458
1,823
15,361
4,364
5,950
53,956
2018
a
28,823
1,705
14,330
4,486
5,498
54,842
2019
a
34,195
1,289
15,771
5,126
5,971
62,352
2020
a
71,733
3,203
7,298
8,223
6,890
97,347
2021
a
59,565
3,207
12,079
7,485
6,995
89,331
2022
a,f
56,178
2,662
a
Preliminary
b
Unknown harvests (Mexico) were assumed to be 10% of harvests in the U.S. and Canada.
c
Unretrieved kill as reported by hunters is used for the Central Flyway; for the remainder of harvest areas, it is assumed to be
20% of retrieved harvests.
d
There is no estimate available for AK in that year.
e
Estimates (except Canada) biased low because of HIP sampling issues that led to estimates of zero harvest in SD and AK.
f
Estimates for Canada, Mexico, North America, and Total not calculated because data for Canada was not available.
43
Table 7B. Decadal and long-term annual averages of annual sport hunting mortality estimates for the
Mid-Continent Population of sandhill cranes in North America.
Year
Central
Flyway
Other Survey
Area Total
Canada
Mexico
Unretrieved
No. America
Total
1975-79
9,913
592
2,663
1,317
2,673
17,158
1980-89
11,688
1,171
7,051
1,991
3,860
25,761
1990-99
17,206
1,036
7,600
2,576
4,231
32,565
2000-09
16,925
1,037
10,735
2,870
4,504
36,070
2010-19
21,044
1,748
13,883
3,667
5,171
45,513
2020-22
62,492
3,024
1975-2022
18,868
1,296
9,051
2,837
4,360
35,565
Table 8. Spring population indices for Rocky Mountain sandhill cranes in the San Luis Valley, Colorado,
1984-96.
Year
Raw
Count
Adjusted for
Estimation
Bias
a
Adjusted by
Removal of
Lesser sandhill
Cranes
b
Other
Areas
Index
Survey
Condition
1984
10,962
14,488
13,562
550
14,112
POOR
1985
18,393
21,773
20,382
0
20,382
GOOD
1986
14,031
14,031
13,135
20
13,155
POOR
1987
13,561
15,661
14,660
0
14,660
POOR
1988
17,510
17,510
16,381
22
16,403
POOR
1989
17,302
18,389
17,004
0
17,004
GOOD
1990
20,851
24,593
21,221
275
21,496
GOOD
1991
19,990
18,405
16,045
175
16,220
GOOD
1992
23,516
23,516
19,999
9
20,008
GROUND
1993
17,576
17,576
16,478
1,260
17,738
POOR
1994
17,229
16,036
15,063
203
15,266
FAIR
1995
25,276
23,390
20,229
0
20,229
GOOD
1996
23,019
26,379
22,737
1,010
23,747
GOOD
a
Raw estimate adjusted by photography for estimation bias.
b
Population estimate adjusted to remove the number of lesser sandhill cranes (non-RMP cranes).
44
Table 9. Fall pre-migration population indices for Rocky Mountain sandhill cranes.
Year
UT
CO
ID
WY
MT
TOTAL
3-Year AVG
1987
1,578
1,443
10,686
2,327
1,447
17,481
1992
2,810
3,181
5,801
2,248
5,264
19,304
1995
1,528
2,284
6,864
1,671
3,681
16,028
1996
1,849
1,255
8,334
2,526
2,974
16,938
1997
a,b
2,450
1,604
8,132
2,255
3,595
18,036
17,001
1998
2,185
1,273
8,067
3,162
3,415
18,102
17,692
1999
2,292
1,102
8,761
4,205
3,141
19,501
18,546
2000
2,416
749
9,337
3,890
3,598
19,990
19,198
2001
1,522
666
7,160
2,626
4,585
16,559
18,683
2002
1,869
1,355
7,698
3,038
4,843
18,803
18,451
2003
2,546
745
7,822
3,446
4,964
19,523
18,295
2004
2,239
1,410
7,152
3,072
4,637
18,510
18,945
2005
2,646
1,052
7,668
3,911
5,588
20,865
19,633
2006
c
NS
19,633
2007
d
2,401
1,743
8,262
3,907
6,509
22,822
20,732
2008
e
3,708
1,080
6,123
3,826
6,419
21,156
21,614
2009
2,283
1,162
6,934
3,613
6,329
20,321
21,433
2010
3,242
985
5,776
3,726
7,335
21,064
20,847
2011
1,498
1,347
5,029
2,978
6,642
17,494
19,626
2012
2,109
413
3,432
3,587
5,876
15,417
17,992
2013
2,732
1,594
5,228
3,588
7,218
20,360
17,757
2014
2,783
1,258
6,064
3,008
6,555
19,668
18,482
2015
3,698
1,089
6,454
3,596
9,493
24,330
21,453
2016
f
3,298
1,135
5,445
4,879
7,507
22,264
22,087
2017
2,994
1,658
4,066
3,725
7,149
19,592
22,062
2018
2,770
1,908
4,469
5,101
7,553
21,801
21,219
2019
3,106
1,879
4,428
4,366
7,511
21,290
20,894
2020
3,222
1,446
5,096
6,608
9,264
25,636
22,909
2021
3,889
3,141
3,091
6,059
7,783
23,963
23,630
2022
2,330
1,526
3,957
3,975
6,844
18,632
22,744
a
Incomplete survey efforts in years prior might have resulted in lower estimates; the official count begins in 1997.
b
In October 1997, a special survey was also conducted in the SLV, Colorado and other areas, which resulted in a total
of 27,090 Rocky Mountain and Mid-Continent cranes being counted.
c
In 2006, the survey was not conducted due to mechanical issues with the survey plane. The 3-yr Avg for 2006 is calculated using
2003-05.
d
The 3-yr average for 2007 was calculated using 2004, 2005, and 2007 because there was no survey in 2006.
e
The 3-yr average for 2008 was calculated using 2005, 2007, and 2008 because there was no survey in 2006.
f
Beginning 1n 2016 Wyoming added six new survey areas as allowed in the management plan.
45
Table 10A. Estimated retrieved harvests of the Rocky Mountain Population of sandhill cranes.
Year
UT
NM
AZ
WY
MT
ID
Total
1981
20
20
1982
9
143
152
1983
35
154
189
1984
33
101
134
1985
40
138
178
1986
23
195
218
1987
60
190
250
1988
310
40
128
478
1989
54
483
51
125
713
1990
35
79
9
58
181
1991
48
47
44
101
240
1992
0
147
39
168
42
396
1993
28
297
61
115
45
546
1994
34
416
27
150
40
667
1995
27
270
33
77
41
448
1996
32
236
27
84
49
20
448
1997
30
114
22
82
62
136
446
1998
34
180
37
93
59
135
538
1999
54
198
21
124
71
190
658
a
2000
69
257
37
163
91
193
810
b
2001
77
288
26
142
87
278
898
2002
60
164
42
132
51
194
643
2003
57
169
34
72
50
146
528
2004
53
189
35
124
51
142
594
2005
62
236
50
116
49
189
702
2006
87
327
10
194
54
235
907
2007
103
276
43
138
73
187
820
2008
101
379
24
162
85
185
936
2009
149
603
67
195
124
254
1,392
2010
190
547
56
182
108
253
1,336
2011
c
149
522
37
166
90
293
1,257
2012
c
91
417
85
134
129
275
1,131
2013
c
95
241
38
74
94
135
677
2014
73
183
20
94
121
134
625
2015
86
145
67
104
137
166
705
2016
72
453
74
158
140
258
1,155
2017
189
395
68
193
150
198
1,193
2018
203
623
102
189
154
253
1,524
2019
130
456
38
141
179
166
1,110
2020
137
160
61
114
151
172
795
2021
200
572
63
194
129
277
1,435
2022
223
656
69
227
138
273
1,586
a
RMP sandhill cranes (40) were also taken as part of research project in the San Luis Valley, CO
b
RMP sandfill cranes (20) were also taken as part of research project in the San Luis Valley, CO
c
Harvest includes crippling loss.
46
Table 10B. Decadal and long-term annual averages of retrieved harvests of the Rocky Mountain
Population of sandhill cranes.
Years
UT
NM
AZ
WY
MT
ID
Total
1981-89
397
35
147
259
1990-99
32
198
32
105
51
120
434
2000-09
82
289
37
144
72
200
824
2010-19
128
398
59
144
130
213
1,071
2020-22
187
463
64
178
139
241
1,272
1981-2022
89
315
42
137
92
198
705
Table 11. Winter counts of Lower Colorado River Valley Population of sandhill cranes in Arizona and
California.
Year
Cibola
National
Wildlife
Refuge
Colorado River
Indian Tribe
Salton Sea
National
Wildlife
Refuge
Gila
River
Total
3-Year AVG
1998
775
596
351
178
1,900
1999
1,200
511
325
163
2,199
2000
820
1,259
235
252
2,566
2,222
2001
961
952
350
134
2,397
2,387
2002
1,003
168
417
52
1,640
2,201
2003
1,200
455
430
0
2,085
2,041
2004
1,341
354
521
312
2,528
2,084
2005
1,513
457
476
191
2,637
2,417
2006
1,141
673
493
360
2,667
2,611
2007
2,322
809
295
450
3,876
3,060
2008
a
115
NS
687
413
1,215
3,060
2009
b
289
1,216
603
293
2,401
2,981
2010
c
266
729
904
365
2,264
2,847
2011
553
636
899
327
2,415
2,360
2012
1,097
474
924
151
2,646
2,442
2013
1,629
344
671
434
3,078
2,713
2014
1,981
591
641
140
3,353
3,026
2015
676
720
688
452
2,536
2,989
2016
631
631
862
292
2,416
2,768
2017
940
636
819
321
2,716
2,556
2018
1,076
330
775
215
2,396
2,509
2019
1,171
192
1,062
497
2,922
2,678
2020
1,497
20
1,105
319
2,941
2,753
2021
4,812
0
954
117
5,883
3,915
2022
941
1,248
1,226
372
3,787
4,204
2023
864
2186
1532
137
4,719
4,796
NS = No survey was conducted.
a
In 2008, the survey was not complete. The 3-YR average for that year was calculated using 2005-07.
b
In 2009, the 3-YR average was calculated with 2006, 2007 and 2009 due to an incomplete survey in 2008.
c
In 2010, the 3-YR average was calculated with 2007, 2009, and 2010 due to an incomplete survey in 2008.
47
Table 12. Fall abundance index for Eastern Population of sandhill cranes.
Year
Abundance
3-YR Average
1979
14,385
1980
15,808
1981
11,943
14,045
1982
13,879
13,877
1983
16,148
13,990
1984
16,363
15,463
1985
16,170
16,227
1986
17,043
16,525
1987
22,342
18,518
1988
26,086
21,824
1989
22,785
23,738
1990
23,792
24,221
1991
24,685
23,754
1992
26,656
25,044
1993
26,187
25,843
1994
26,783
26,542
1995
33,774
28,915
1996
29,753
30,103
1997
27,641
30,389
1998
37,827
31,740
1999
33,583
33,017
2000
33,105
34,838
2001
a
NS
34,838
2002
b
31,575
32,754
2003
c
29,300
31,327
2004
28,822
29,899
2005
37,708
31,943
2006
37,529
34,686
2007
37,943
37,727
2008
44,110
39,861
2009
60,028
47,360
2010
49,647
51,262
2011
76,028
61,901
2012
86,989
70,888
2013
64,213
75,743
2014
82,694
77,965
2015
94,676
80,528
2016
70,858
82,743
2017
69,989
78,508
2018
97,073
79,307
2019
89,504
85,522
2020
94,961
93,846
2021
d
90,029
91,498
2022
107,164
97,385
NS = No survey conducted
a
In 2001, the survey was not conducted. The 3-YR average for that year was calculated using data from 1998-2000.
b
In 2002, the 3-YR average was calculated with 1999, 2000 and 2002 since the survey was not conducted in 2001.
c
In 2003, the 3-YR average was calculated with 2000, 2002 and 2003 since the survey was not conducted in 2001.
d
Illinois and Maryland began participating in the survey in 2021.
48
Table 13. Season dates (month/day) for the hunting of Eastern Population sandhill cranes.
Year
Kentucky
Tennessee
Alabama
2011
12/17-01/15
No Season
No Season
2012
12/15-01/13
No Season
No Season
2013
12/14-01/12
11/28-01/01
No Season
2014
12/13-01/11
11/22-11/23; 11/29-01/01
No Season
2015
12/12-01/10
11/28-11/29; 12/05-01/01
No Season
2016
12/17-01/15
12/03-01/12; 01-16-01/29
No Season
2017
12/16-01/14
12/02-01/28
a
No Season
2018
12/03-01/27
12/01-01/27
b
No Season
2019
12/02-01/26
12/07-01/27
c
12/03-01/05; 01/16-01/31
2020
12/07-01/31
12/05-01/31
d
12/04-01/03; 01/11-01/31
2021
12/07-01/31
12/04-01/30
d
12/02-01/09; 01/17-01/31
2022
12/07-01/31
12/03-01/31
d
12/03-01/08; 01/16-01/31
a In the Southeast Zone, the season was closed from 01/12-01/14.
b In the Southeast Zone, the season was closed from 01/18-01/20.
c In the Southeast Zone, the season was closed from 01/17-01/19.
4 In the Southeast Zone, the season was closed from 01/15-01/17.
e In the Southeast Zone, the season was closed from 01/14-01/16.
Table 14. Estimated harvest and number of permits sold for Eastern Population sandhill cranes.
Year
KY
Harvest
KY Tags
Issued
a
TN
Harvest
TN Tags
Issued
a
AL
Harvest
AL Tags
Issued
a
Total
Harvest
Total
Permits
Issued
2011
50
534
No Season
No Season
No Season
No Season
50
534
2012
92
570
No Season
No Season
No Season
No Season
92
570
2013
87
570
350
1,200
No Season
No Season
437
1,770
2014
96
704
393
1,200
No Season
No Season
489
1,904
2015
75
694
161
1,200
No Season
No Season
236
1,894
2016
171
672
586
1,200
No Season
No Season
757
1,872
2017
119
660
830
2,319
No Season
No Season
949
2,979
2018
60
1,432
555
2,711
No Season
No Season
615
4,143
2019
96
1,237
746
2,958
291
1,200
1,133
5,395
2020
65
1,035
630
2,700
391
1,200
1,086
4,935
2021
117
1,029
484
2,500
234
1,200
835
4,729
2022
180
1,148
640
2,500
265
1,200
1,085
4,848
Average
101
857
538
1,999
295
1,200
607
2,793
a
Each tag allows a hunter to take one crane.
49
Figure 1. Primary wintering and breeding range and the approximate migration corridor of Mid-
Continent sandhill cranes (based on figures in Tacha et al. 1994, Krapu et al. 2011).
50
Figure 2. Approximate range of the Rocky Mountain Population of Greater sandhill Cranes
(Tacha et al. 1994, Drewien et al. 1996).
51
Figure 3. Approximate range of the Lower Colorado River Valley Population of Greater sandhill
Cranes (based on Pacific Flyway Council [1995] and recent satellite telemetry information [D.
Collins and K. Kruse, U.S. Fish and Wildlife Service, unpublished data]).
52
Figure 4. Approximate range of Eastern Population sandhill cranes based on various data
sources including satellite telemetry data, breeding bird atlas records, and unpublished location
information from knowledgeable individuals.
53
Figure 5. Areas open to the hunting of Mid-continent sandhill cranes by Federal frameworks in
the Central Flyway states, 2021-22.
54
Figure 6. Spring population indices for Mid-Continent sandhill cranes on the Central Platte River
Valley, Nebraska. Survey was not conducted in 2020.
Figure 7. Photo-corrected spring population estimates (solid line) and the 95% confidence
intervals (dashed lines) for Mid-Continent sandhill cranes on the Central Platte River Valley,
Nebraska. The survey was not conducted in 2020.
55
Figure 8. Annual and three-year average photo-corrected ocular transect spring population
indices and population objective thresholds for Mid-Continent sandhill cranes.
Figure 9. Active Mid-Continent sandhill crane hunters in the U.S. portion of the Central Flyway.
56
Figure 10. Crippling-loss rate (number lost/[number retrieved + lost]) of Mid-Continent sandhill
cranes in the U.S. portion of the Central Flyway.
Figure 11. Average number of hunting days afield reported by active Mid-Continent sandhill
crane hunters in the U.S. portion of the Central Flyway.
57
Figure 12. Seasonal bag per Mid-Continent sandhill crane hunter in the U.S. portion of the
Central Flyway.
Figure 13. Estimated hunting mortality (retrieved plus unretrieved) of Mid-Continent sandhill
cranes in the U.S. portion of the Central Flyway.
58
Figure 14. Estimated hunting mortality (retrieved plus unretrieved) of Mid-Continent sandhill
cranes in North America. Data unavailable for Canada and Mexico for 2022.
Figure 15. Relationship between indices of abundance and harvest of Mid-Continent sandhill
cranes. Data unavailable for Canada and Mexico for 2022.
59
Figure 16. Estimated harvest of Rocky Mountain Population sandhill cranes.
Figure 17. Abundance indices for the Rocky Mountain Population of sandhill cranes.
60
Figure 18. Annual and three-year average of fall pre-migration abundance indices for the Rocky
Mountain Population of sandhill cranes.
Figure 19. Annual indices for recruitment (% juveniles) of the Rocky Mountain Population of
sandhill cranes. Solid line indicates the long-term (1972-2021) average of 8.2.
61
Figure 20. Annual and three-year average of winter counts of the Lower Colorado River Valley
Population of sandhill cranes in Arizona and California.
Figure 21. Annual and three-year average of fall counts of the Eastern Population of sandhill
cranes.
U.S. Fish and Wildlife Service
Division of Migratory Bird Management
Branch of A
ssessment
and Decision Support
11510 American Holly Drive
Laurel, Maryland
20708-4002
http://www.fws.gov
August
2023
For State Transfer Relay Service: TTY/ Voice: 711